Table 1.

Molecular variation in CO1 in six species of Helictopleurus and in Nanos clypeatus (Canthonini)

Species	n	bp	V%	Pi%	S%	Diet	Habitat
Helictopleurus marsyas	44	688	0.44	0.15	0.29	Cattle dung	Open area
Helictopleurus neoamplicollis	112	688	1.02	0.15	0.87	Cattle dung	Open area
Helictopleurus quadripunctatus	54	688	5.81	3.34	2.47	Cattle and human dung, carrion	Open area and dry forest
Helictopleurus nicollei	5	688	0.00	0.00	0.00	Carrion and other dung	Wet forest
Helictopleurus perrieri	11	688	5.09	3.63	1.45	Carrion and other dung	Dry forest
Helictopleurus unifasciatus	29	688	5.96	3.63	2.33	Carrion and other dung	Dry forest
Nanos clypeatus	54	771	10.89	9.21	1.69	Carrion and other dung	Wet forest

Helictopleurus littoralis has been combined here with Helictopleurus neoamplicollis.

V%, Pi% and S% are the percentages of variable, parsimony informative and singleton sites, respectively.