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. Author manuscript; available in PMC: 2009 Sep 3.
Published in final edited form as: Physiol Behav. 2008 May 13;95(1-2):93–100. doi: 10.1016/j.physbeh.2008.05.002

Table 2.

Observed function of facial muscles during intramuscular stimulation in the rhesus macaque (current study) compared to humans and chimpanzees [14].

Muscle Rhesus Human Chimpanzee
Frontalis (inner) Elevates the medial portion of the brow (AU1: inner brow raiser) Elevates the medial portion of the brow (AU1: inner brow raiser) Elevates the medial portion of the brow (AU1: inner brow raiser)
Frontalis (outer) Elevates the lateral portion of the brow (AU2: outer brow raiser) Elevates the lateral portion of the brow (AU2: outer brow raiser) Elevates the lateral portion of the brow (AU2: outer brow raiser)
Corrugator supercilli Lowers the brow inferiorly with slight medial contraction (corrugation) (part of AU4: brow lowerer) Draws the brow medially and superiorly (AU1+AU4: inner brow raiser + brow lowerer) Not stimulated (but likely present)
Depressor supercilli Not present Depresses the medial portion of the brow (part of AU4: brow lowerer) Not stimulated (but may be present)
Procerus Depresses the medial portion of the brow (part of AU4: brow lowerer) Depresses the medial portion of the brow (part of AU4: brow lowerer) Depresses the medial portion of the brow (part of AU4: brow lowerer)
Orbicularis oculi Inferior portion elevates infraorbital triangle (or equivalent area) superiorly and medialwards Elevates the infraorbital triangle (cheek) superiorly and medialwards (AU6: cheek raiser) Inferior portion elevates infraorbital triangle (or equivalent area) superiorly and medialwards
Nasalis Wrinkles skin on bridge of nose (part of AU9) Wrinkles skin on bridge of nose (part of AU9) Not stimulated
Levator labii superioris alaeque nasi Wrinkles the skin alongside the nose (AU9: nose wrinkler) Wrinkles the skin alongside the nose (AU9: nose wrinkler) Wrinkles the skin alongside the nose (AU9: nose wrinkler)
Levator labii superioris Elevates upper lip (AU10: upper lip raiser) Not stimulated (but present). Elevates upper lip (AU10: upper lip raiser)
Zygomatic minor Not stimulated (but may be present) AU11 (nasolabial furrow deepener) Not stimulated (but may be present)
Zygomatic major Elevates lip corners superiorly and draws lip corners laterally, increasing angle of the mouth (AU12: lip corner puller) Elevates lip corners superiorly and draws lip corners laterally, increasing angle of the mouth (AU12: lip corner puller) Elevates lip corners superiorly and draws lip corners laterally, increasing angle of the mouth (AU12: lip corner puller)
Buccinator Not stimulated (but present) Not stimulated (but present) Not stimulated (but present)
Depressor anguli oris Depresses lip corners (AU15: lipcorner depressor) Depresses lip corners (AU15: lip corner depressor) Depresses lip corners (AU15: lip corner depressor)
Depressor labii inferioris Depresses medial portion of lower lip (AU16: lower lip depressor) Depresses medial portion of lower lip (AU16: lower lip depressor) Depresses medial portion of lower lip (AU16: lower lip depressor)
Mentalis Pushes skin of the chin boss superiorly (AU17: chin raiser) Pushes skin of the chin boss superiorly (AU17: chin raiser) Pushes skin of the chin boss superiorly (AU17: chin raiser)
Orbicularis oris Reduces lip aperture and protrudes lips causing pursing (AU18: lip pucker) Tightens lip margins (AU23: lip tightener). Other FACS movements not stimulated (AU18:lip pucker, AU22: lip funneler,AU24: lip presser; AU28: lip suck) Reduces lip aperture and funnels/protrudes lips (AU22: lip funneller)