Submotif song features and permutations. Example spectrograms from excerpts of three test exemplars where ordering of the submotif features, denoted by the black overlaid boxes, is (a) unpermuted (i.e., naturally ordered), (b) permuted within the temporal boundaries of their original motif, or (c) permuted over the entire exemplar. The submotif features for the second and third motifs in (a) are labeled with letters, so that their permuted positions in (b) and (c) can be more easily seen. Because only an excerpt of the entire stimulus exemplar is shown, not all of the submotif features labeled in (a) appear in (c).