Pressure-volume relationship of a lymphatic segment. Stroke volume, the difference between end-diastolic volume (Ved) and end-systolic volume (Ves), is determined using inlet pressure (Pin), outlet pressure (Pout), slope of end-systolic pressure-volume relationship (Emax), slope of end-diastolic pressure volume relationship (Emin), and dead volume [i.e., volume at 0 pressure (V0)].