Fig. 1.

Three representative circuit units of the model. Each unit includes eight neuronal elements: PR, photoreceptor; HC, horizontal cell; on-BC, on-type bipolar cell; off-BC, off-type bipolar cell, on-AC, on-type amacrine cell; off-AC, off-type amacrine cell; wf-AC, wide-field on-off amacrine cell; GC, on-off ganglion cell. Excitatory and inhibitory synaptic connections are represented with white and black triangles, respectively. Gap junctions between HC elements are indicated by dotted horizontal lines. Wide-field lateral connections are implemented between wf-AC and GC. Outer retinal units (i.e., PR, HC, on-BC, and off-BC) are enclosed with a gray dashed box constitute an outer retinal computational unit. Note that only one computational circuit unit is fully illustrated, remote units are partially shown to highlight distal connections