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MGU morphology of mature pollen in +/spp-2. The nuclei of mature pollen grains from +/spp-2 plants were visualized by DAPI staining. Wild-type pollen grains (A and D) show a typical arrangement of the MGU, consisting of a well-centered vegetative nucleus (white arrowhead) and two sperm cells (black arrowheads). Pollen from +/spp-2 plants shows an abnormal gum-like (B and E) or mud-like (C and F) MGU morphology characteristic of the gum and mud mutants (Lalanne and Twell, 2002).
