Distribution of style spreading in Phyllantheae. Species pollinated by Epicephala have reduced styles that are medially fused, whereas non-Epicephala-pollinated species have horizontally spread, bifid styles. Filled and open boxes indicate species with and without associations with Epicephala, respectively. Ecological data were not available for species with asterisks. Female flowers are drawn for Phyllanthus marojejiensis, G. acuminatum, Sauropus brevipes and Flueggea suffruticosa (from left to right). Error bars, ±1 s.e.