TABLE 2.

Genetic correlations between males and females (diagonal), between traits in males (above diagonal), and between traits in females (below diagonal), across all ages assuming no genotype-by-environment interaction

	Horn length	Body weight	Parasite load
Horn length	0.223 ± 0.201	0.541 ± 0.143**	0.151 ± 0.178
Body weight	0.097 ± 0.168	0.793 ± 0.246**	−0.530 ± 0.210**
Parasite load	0.093 ± 0.202	−0.278 ± 0.169	0.838 ± 0.273***

Estimates and standard error (± SE) were obtained from multivariate animal models (see methods, model 2). Significance from zero was assessed using log-likelihood-ratio tests: *P < 0.050; **P < 0.010; *** P < 0.001.