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. 2009 May 1;5(5):e1000373. doi: 10.1371/journal.pcbi.1000373

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Jehee et al.

This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are properly credited.

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Higher-level coding units attempt to predict the responses of units in the next lower level via feedback connections, while lower-level error detectors signal the difference between the prediction and the actual input. Feedforward connections encode the synaptic weights represented by a matrix . Higher-level units maintain the current estimate of the input signal and convey the top-down prediction to the lower level via feedback connections. Difference detectors compute the difference between current activity and the top-down prediction .

Inline graphic — Higher-level coding units attempt to predict the responses of units in the next lower level via feedback connections, while lower-level error detectors signal the difference between the prediction and the actual input. Feedforward connections encode the synaptic weights represented by a matrix . Higher-level units maintain the current estimate of the input signal and convey the top-down prediction to the lower level via feedback connections. Difference detectors compute the difference between current activity and the top-down prediction .