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. 2008 Nov 10;364(1516):497–501. doi: 10.1098/rstb.2008.0250

Figure 1.

Figure 1

Four stages in a demonstration of countershading, from left to right: (a) a flat expanse of paper; (b) the artistic tradition of shading (or top–down lighting), by which a flat surface takes on the appearance of volume; (c) countershading, by which the undersides of animals are lighter than the surfaces that have greater exposure to sunlight; and (d) the flat expanse of tone that comes from shading being cancelled out by countershading. Author's diagram.