Table 2.

Fruit developmental landmarks.

Fruit Development Landmarks	Days post anthesis	Descriptions of fruit development in tomato
		Fruit growth (Gillaspy et al 1993)	Embryo/seed development
(1) Anthesis	0	Mature ovary, phase I.	Mature gametes. Pollen is shed, which will land on the stigma and germinate. Pollen tubes growth through the style.
(2) Fertilization	1–2	End of phase I, beginning of phase II.	Fusion of sperm and egg nuclei.
(3) 4–16 Cell Stage Embryo	3–6	Phase II and III, cell division and elongation stage.	First embryo divisions.
(4) Globular Stage Embryo	6–10	Phase III, cell expansion stage.	Globular embryo.
(5) Heart Stage Embryo	10–12	Phase III, cell expansion stage.	Heart Stage embryo lasts approximately one day and occurs 10–12 dpa.
(6) Torpedo Stage Embryo	13–16	Phase III, continued fruit enlargement.	Torpedo Stage embryo lasts approximately one day and occurs 13–16 dpa.
(7) Coiled Stage Embryo	20	Phase III, continued fruit enlargement.	Cotyledon expansion and curl as they elongate. Embryo appears physically mature, but the seed is not yet viable.
	20–28		Seed maturation period
(8) Seed germination	29–31	The fruit has reached the mature green stage. Fruit becomes sensitive to ethylene.	Seeds are becoming viable for germination.
(9) Fruit ripening	33–40	Ripening starts at the onset of the breaker stage. Changes in pigmentation are visible.	After ripening of seed.
(10) Ripe Fruit	40	Red ripe stage of tomato.

Timing of the fruit landmarks in S. pimpinellifolium LA1589.