Examples of combined phylogeographic, bioacoustic, and morphological evidence used to classify divergent mitochondrial lineages as CCS or DCL. (A) Boophis majori and B. aff. majori are sympatric CCS; despite their only low genetic divergence, they have distinct and constant differences in tadpole morphology and qualitative differences in advertisement calls, without signal of genetic admixture. (B and C) B. luteus (B) and Guibemantis liber (C) consist of deeply divergent genealogical lineages classified as DCL, because the call differences only affect quantitative parameters such as note-repetition rate in B. luteus, and there are no morphological or ecological differences between the populations. Furthermore, genetic admixture of the lineages was detected in G. liber at one locality.