Fig. 3.

Graphical depiction of how floral optimality and precision are manifested on the pollinator. The distribution of pollen on the bee is determined by the population variance in gland–anther distance (GAD) and the distribution of stigma contacts is determined by the population variance in gland–stigma distance (GSD). The departure from optimum (the difference between the means of the two distributions) reflects, in turn, the difference between population mean GAD and population mean GSD. The bee is a male Euglossa viridissima, pollinator of Dalechampia spathulata, the females of which pollinate D. scandens.