Table 3.

Measurements and landmarks used to calculate male and female inaccuracies and their components in the four study systems

Study system	Landmark	Male accuracy trait	Male fitness optimum	Female accuracy trait	Female fitness optimum	Calculation of mean-scaled accuracy index
Linum suffruticosum (class 2)	Inner face of petal	Petal–anther gap (thrum)	Petal–stigma gap (pin)	Petal-stigma gap (pin)	Petal–anther gap (thrum)	[(mean gappin – mean gapthrum)2 + VARgap-pin + VARgap-thrum]/(trait mean)2
Dalechampia spp. (class 3)	Resin gland	Gland–anther distance (GAD)	Population mean gland–stigma distance (GSD)	GSD	Population mean GAD	[(mean GSD –mean GAD)2 +VARGSD + VARGAD]/(trait mean)2
Collinsia spp. (class 4)	Throat of floral tube	Throat–anther distance (TAD)	Population mean throat–stigma distance (TSD)	TSD	Population mean TAD	[(mean TSD – mean TAD)2 + VARTSD +VARTAD]/(trait mean)2
Stylidium spp. (class 5): column length	Sensitive trigger point on column	Trigger–anther distance with column sprung (‘length’)	Population mean trigger–stigma distance (column sprung)	Trigger–stigma distance (column sprung)	Population mean trigger–anther distance (column sprung)	[(mean lengthmale – mean lengthfemale)2 + VARlength-male + VARlength-female)]/(trait mean)2
Stylidium spp. (class 5): column reach	Sensitive trigger point on column	Location of pollen on artificial pollinator (‘reach’)	Population mean column reach	Column reach	Population mean location of pollen on pollinator	[(mean reachmale – mean reachfemale)2 + VARreach-male + VARreach-female]/(trait mean)2