Whiteness in cuttlefish is created by scattering of light from leucophores. (a) Disruptive pattern, (b) zebra pattern, (c) close-up of white square, (d) close-up of zebra pattern, (e) close-up of white finspot, created by leucophores. Note the chromatophores in the superficial layer; chromatophores can modulate whiteness. (f) Scanning electron micrograph of leucophores showing spherical arrangement of leucosomes (courtesy of Alan Kuzirian).