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. 2009 Mar 11;103(7):1049–1064. doi: 10.1093/aob/mcp048

Table 1.

Habit and habitat details of the woody species studied

Species Habit Habitat and distribution
Spermacoceae
Arcytophyllum lavarum Subshrub Volcanic mountains at high altitude; Costa Rica to Panama
Arcytophyllum setosum Shrub Grassland above timberline; western South America
Arcytophyllum thymifolium Subshrub or shrub Grassland above timberline; Colombia to Peru
Diodella sarmentosa Straggling, scrambling or climbing, often perennial herb Large ecological amplitude, including evergreen forest, riverine vegetation, bush land, rocky places; widespread in the tropics
Emmeorhiza umbellata Climbing, perennial herb Tropical forest; South America
Gomphocalyx herniarioides Prostrate or decumbent herb with well-developed, often woody taproot Sandy soils of thorn forest, dunes and beaches; Madagascar
Hedyotis flavescens Shrub Upper montane zone on open waterlogged sites; Sri Lanka
Hedyotis fruticosa Shrub or small tree to 4 m Open vegetation, rocky areas on mountain tops; southern India, Sri Lanka, Myanmar
Hedyotis lessertiana Shrub or small tree to 4 m Upper montane scrubland; Sri Lanka
Hedyotistrichoglossa Woody herb of suffrutex with slightly woody stems and taproot Eastern humid forest; Madagascar
Kadua cordata Subshrub or shrub Mesic to wet forests; Hawaiian islands
Lathraeocarpa acicularis Subshrub with woody stems and taproot Sandy soils in dunes close to the sea; Madagascar
Mitracarpus frigidus Subshrub or shrub Scrubland, grassland, also in disturbed areas; tropical America
Nesohedyotis arborea Tree up to 7 m Tree fern thicket; St. Helena island
Oldenlandiaambovombensis Shrub Dry spiny forest on limestone plateau; south-western Madagascar
Oldenlandiahumbertii Subshrub Dry spiny forest on limestone plateau; south-western Madagascar
Spermacoce bangweolensis Prostrate or suberect subshrub with woody taproot Sandy soil bordering Lake Bangweulu; Zambia
Spermacoce manikensis Geoxylic herb with woody taproot and plant base High plateaus on Kalahari sand; Zambia and D.R. Congo
Spermacoce macrocephala Subshrub Savanna areas on river banks; Colombia and Venezuela
Spermacoce occidentalis Annual or more often perennial spreading or erect woody herb Open woodland on sandy soil or dune vegetation close to sea; Australia
Spermacoce verticillata Woody herb or (sub)shrub Savanna, scrubland and disturbed areas; tropical and subtropical America, introduced elsewhere
Knoxieae
Carphalea kirondron Shrub or tree up to 10 m Mostly at edges of dry, deciduous forest on laterite or calcareous sand; Madagascar
Dirichletia virgata Shrub Forest and scrubland; Socotra, Republic of Yemen
Otiophora rupicola Subshrub Rocky grassland at high altitude; Burundi
Otomeria micrantha Herb, often with woody base Savanna, clearings in forest, along roadsides; Nigeria to west-central tropical Africa
Pentanisia schweinfurthii Geoxylic herb with woody taproot and plant base Submontane and lower grassland, also in clearings of woodland; tropical Africa
Pentas zanzibarica Woody herb or shrub with a woody rootstock Forest edge, scrubland and grassland; Uganda to Mozambique
Phyllopentas schimperiana Woody herb or shrub Forest edge and scrubland, at high altitudes; tropical Africa
Triainolepis polyneura Shrub Deciduous and semi-deciduous forest; Madagascar