θ1	orientation preference of point 1;
θ2	orientation preference of point 2;
s1	orientation selectivity of point 1;
s2	orientation selectivity of point 2;
r	cortical distance between points 1 and 2;
φ	angle of the line from point 1 to point 2;
α = θ2 − θ1	relative orientation preference of the two points;
β = φ − θ1	angular position of the second point relative to the orientation preference of the first point.