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Arrangement of the active site of Aa_Cel9A in complex with substrate (cellotetraose). Glu515 acts as a proton donor and Asp143 and Asp146 are thought to deprotonate the water (W548) involved in the nucleophilic attack. The modeling analyses and the docking of the cellotetraose molecule into the active site of Aa_Cel9A were performed based on the inactive mutant of the Ct_CbhA–cellotetraose complex (Schubot et al., 2004 ▶).
