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Mean fecal progesterone and glucocorticoid metabolite concentrations were not detectably related to changes in population density. Relative population density was measured as a population's current size divided by the long-term mean size of that population, so that a value of one indicates a population at its long-term mean density. (A) Fecal progesterone concentration. (B) Winter-long mean fGCM concentration. (C) Early winter mean fGCM concentration.
