One of 725 most-parsimonious trees (MPTs) of all molecular data combined (DELTRAN optimization). Branch lengths are above the branches, and bootstrap percentages are below the branches. Branches that collapse in the strict consensus tree are indicated by closed circles. Taxa missing a leaf abscission layer are enclosed within the box. Fruit ornamentation is also shown with character states [glabrous (o) vs. setose (*)] on branch terminals, and hash-marks in the tree represent apomorphies. Traditional sectional placement for all species is listed by abbreviation (sections Dichaea, DICH; Dichaeastrum, ASTR; Dichaeopsis, OPSI; Pseudodichaea, PSEU); present circumscription is listed on the far right.