Table 1.

Analysis of variance format for frequencies of allele A₁.

Source	d.f.	Mean Square	Expected Mean Square
Among Populations	r − 1	${\text{MSA}}_1=\frac{{\sum }_{i=1}^r{n}_i{({\stackrel{\sim }{p}}_{i1}-{\overline{p}}_1)}^2}{r-1}$	p1(1 − p1)(l − θ) +ncθp1(1 − p1)
Within Populations	${\sum }_{i=1}^r(n_i-1)$	${\text{MSW}}_1=\frac{{\sum }_{i=1}^r{\sum }_{j=1}^{n_i}{\stackrel{\sim }{p}}_{i1}(1-{\stackrel{\sim }{p}}_{i1})}{{\sum }_{i=1}^r(n_i-1)}$	p1(l − p1)(l − θ)

$n_c=\frac{1}{r-1}\left({\sum }_{i=1}^r{n}_i-\frac{{\sum }_{i=1}^r{n}_i^2}{{\sum }_{i=1}^r{n}_i}\right)$