Flow chart for simulation used. Beginning with cells randomly distributed with zero velocity and acceleration, cells move and interact under the influence of parameters defining viscosities (hfluid and hcytoplasm), interaction strengths (k1AA, k1BB, k1AB, and k0), and computational features such as the size of the computational domain and time step (Δt). Cells that are close to one another (i.e., within dmax) are attracted or repelled with prescribed heterotypic (k1AA, k1BB) or heterotypic (k1AB) interaction strengths, and cells that overlap are repelled hydrostatically with a different strength (k0).