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| Properties of the substratum | Properties of the bulk fluid | Properties of the cell |
|---|---|---|
| Texture or roughness | Flow velocity | Cell surface hydrophobicity |
| Hydrophobicity | pH | Fimbriae |
| Conditioning film | Temperature | Flagella |
| Cations | Extracellular polymeric substances | |
| Presence of antimicrobial agents |
