TABLE 1.

Parameter notations with their corresponding scaled name

Unscaled	Scaled	Description
u		Per-nucleotide, per-generation mutation rate
g		Generation time in years
Ne		Effective population size of the human population
THC	τHC = THCu/2/Ne/g	Human–chimpanzee speciation time
THCG	τHCG = THCGu/2/Ne/g	HC–gorilla speciation time
NHC	θHC = NHCu/2/Ne	HC effective ancestral population size
NHCG	θHCG = NHCGu/2/Ne	HCG effective ancestral population size
r	ρ = r/u	Per nucleotide, per generation, recombination rate (per mutation for the scaled version)

HC and HCG: human–chimpanzee and human–chimpanzee–gorilla ancestral populations.