Collage depicting wide variation in size and shape of tomato fruit. (A) The large-fruited tomato ‘Giant Heirloom’ common to modern agriculture (right) and the typical fruit of a related wild species (L. pimpinellifolium). (B) The range of fruit size and shape variation in tomato. (C) Cross-section of the fruit from a plant homozygous for a mutation at the fasciated locus causing multiple locules. (D) Alternate allele of fasciated associated with unfused carpels. (E) Fruit from ‘Long John’, which carries mutations at both the sun and ovate loci causing extremely long, narrow fruit. (F) Bell pepper-type fruit produced by ‘Yellow Stuffer’. (G) Fruit from two different cultivars homozygous for a mutation at the ovate locus. In the variety on the left, the ovate mutation results in the production of fruit that are both elongated and constricted at the stem end of the fruit (hence, the pear shape). However, in the processing variety on the right, the ovate mutation causes elongated fruit but has a much reduced effect on neck constriction. This figure is from Tanksley SD. 2004. The Plant Cell 16: S181–S189.