Pyramiding introgression lines (ILs) into breeding lines to explore the genetic variation in Brix Yield (BY, measured in g m–2). The upper panel shows the BY differences among different genotypes (shown as ▵ % from M82; base line represents M82) and the lower panel shows the combinations of tomato chromosomes 7, 8 and 9 in these genotypes. Yield differences between M82 (red chromosomes 7, 8 and 9), the four tomato tester inbreds (A = testers, pink chromosomes 7, 8 and 9), and their hybrids (B = M82 × testers) result from allelic variation present in the cultivated tomato gene pool. Since M82, the multiple-introgression line (IL789), and their hybrid IL789 × M82 (C = ILH789) differ only in three S. pennellii segments (green chromosomes), any BY difference between them is associated with the exotic allelic variation. The yield of the hybrids of IL789 with the four testers (D = IL789 × testers) results from both cultivated and exotic variation. This figure is from Gur and Zamir (2004) with modification.