Means and s.e. of (A) decomposition rate Kpot, (B) leaf dry mass content (LDMCgreen), (C) leaf tensile strength (LTSgreen) and (D) resistance to fracture (LRFgreen) for species from different successional stages (E, early; I, intermediate; A, advanced succession) at each nitrogen supply (N− and N+ treatments, as indicated). In the first part of each histogram, the mean and s.e. for all species combined (n = 17 for each N supply) are indicated. Results of post hoc tests (Student–Newman–Keuls comparisons) with successional stage as factor are given with Greek letters (F- and P-values are given in Table 3). Means and s.e. in the 12 species from each taxonomic group are shown in the second part of the histograms (separated by the dotted line). Results of post hoc tests with taxonomic group as factor are given with Roman letters (F- and P-values are given in Appendix 2).