Table 1.

Example annotation output

Verb	AGENT	THEME	LOCATION
activated	NifA: Activator	glnAP2: Gene	In Escherichia Coli: Wild_Type_Bacteria
encodes	GlnA: Gene	glutamine synthetase: Enzyme

For each verb, its separate arguments within the sentence are indicated, together with the biological concepts assigned to them. Each argument is also categorized according to the semantic roles assigned.