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Summary of gene-based genome-wide scans of association of RA with accumulations of rare variants. Each gene-based test has been adjusted for: (a) sex and five axes of genetic variation; (b) sex, five axes of genetic variation, and the number of shared epitope alleles to account for the effects of HLA-DRB1. Genes achieving a nominal significance threshold of p < 10-4 are highlighted in red.
