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. 2009 Dec 22;106(52):22139–22144. doi: 10.1073/pnas.0907520106

Fig. 2.

Fig. 2.

Social linkage and social epistasis. (Left) Linkage occurs because Role 1 and Role 2 may be held by the same person or different people. They are linked by a shared role expectation that explicitly associates them. (Right) Epistasis entails that Role 1 and Role 2 are structurally associated because they are both held by Person A.