Table 2.

The proportion of groups (actual number are in parentheses) with relatedness values greater than random (the 95th percentile of randomized values) and matching haplotypes were calculated across social levels for both breeding females and second-tier group matriarchs (the latter allowing analysis at higher social levels only).

	breeding females		matriarchs
social level	r-values greater than random	matching haplotypes	r-values greater than random	matching haplotypes
second	79% (23 of 29)	84% (21 of 23)	n.a.	n.a.
third	42% (5 of 12)	75% (9 of 12)	25% (3 of 12)	83% (10 of 12)
fourth	0% (0 of 5)	20% (1 of 5)	20% (1 of 5)	40% (2 of 5)