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Depletion of intracellular c-di-GMP levels in the R variant prevents biofilm and rugose colony formation. The indicated strains were grown in HIN. (Top) ATCC 27562R carrying pME6041T or pME6041T::yhjH was grown in the presence (+) or absence (−) of l-arabinose. The arrow marks the position of the biofilm ring. (Bottom) Colony morphology of the strains above. Images shown are representative of at least three independent experiments.
