Angle of incidence at 50% wing length plotted against normalized time through the wingbeat (t̂) for the (a) left and (b) right wings of the five hoverflies. The solid lines show the mean instantaneous angle of incidence for each hoverfly (red, H1; green, H2; blue, H3; black, H4; magenta, H5) and the shaded region around those lines displays the s.d. over the wingbeats measured. The vertical solid line indicates the mean timing of the start of the upstroke.