Table 1. Results of PERMANOVA main test.

Source	df	SS	MS	Pseudo-F	P(perm)	Unique perms
Is	5	17329	3465.9	3.8505	0.0001	9893
Ha	1	3247.3	3247.3	3.6076	0.0021	9949
Qu	2	2010.4	1005.2	1.1168	0.3586	9935
IsxHa	6	10202	1700.4	1.8891	0.0037	9901
IsxQu	10	12160	1216	1.3509	0.0663	9869
HaxQu	4	8608.7	2152.2	2.391	0.0018	9917
IsxHaxQu	3	3326.9	1109	1.232	0.2425	9914
Res	26	23403	900.11
Total	61	1.0056E5

Is = Island, Ha = Habitat type (forereef, backreef, lagoon), Qu = Quadrant (NW, SW, NE, SE).