The average attack rate (attacks per second) ±s.e. of territory holders of four Hetaerina spp. to tethered conspecifics and sympatric congeners. Symbols identify species of the intruders (open circles, H. americana; open triangles, H. occisa; filled squares, H. titia; open squares, H. cruentata). Photographs are of (left to right) H. americana, H. occisa, H. titia and H. cruentata. Site codes are given above the corresponding symbols. Vertical dashed lines connect means for tests carried out at the same site.