EVR is expressed in organ AZs. (A-H) The regulatory regions of EVR and HAE direct expression of β-glucuronidase (GUS) in AZs (A-D), in internal tissues of the floral style (E,F), and at the junction between the cauline leaves and the inflorescence stem (G,H). The HAE::GUS marker is also expressed at the bases of the floral pedicels (B, arrow). Within floral AZs (C), the EVR promoter directs GUS expression prior to organ separation (stage 15), during abscission (stage 16), and as the remaining cells form protective scar tissue (stage 17). GUS expression directed by the HAE promoter shows a similar, yet stronger profile in AZs, with an earlier stage of initiation (Jinn et al., 2000).