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. 2010 Apr 29;6(4):e1000758. doi: 10.1371/journal.pcbi.1000758

Figure 2. Elimination of second-order free-riders (non-punishing cooperators) in the spatial public goods game with costly punishment for Inline graphic, Inline graphic, and Inline graphic.

Figure 2

(a) Initially, at time Inline graphic, cooperators (blue), defectors (red), moralists (green) and immoralists (yellow) are uniformly distributed over the spatial lattice. (b) After a short time period (here, at Inline graphic), defectors prevail. (c) After 100 iterations, immoralists have almost disappeared, and cooperators prevail, since cooperators earn high payoffs when organized in clusters. (d) At Inline graphic, there is a segregation of moralists and cooperators, with defectors in between. (e) The evolutionary battle continues between cooperators and defectors on the one hand, and defectors and moralists on the other hand (here at Inline graphic). (f) At Inline graphic, cooperators have been eliminated by defectors, and a small fraction of defectors survives among a large majority of moralists. Interestingly, each strategy (apart from I) has a time period during which it prevails, but only moralists can maintain their majority. While moralists perform poorly in the beginning, they are doing well in the end. We refer to this as the “who laughs last laughs best” effect.