Counts of infectious (I), recovered (R) and susceptible (S) gerbils (with proportions in parentheses), where captures were divided into those from a burrow system known to contain one or more infectious gerbils the previous trapping session (relatively rare) and those from burrow systems where there was no evidence of recent infection (Pearson's χ2 test: P=0·0027 and 1·8×10−5 for sites 1 and 2 respectively)