FIGURE 3. Crystal structure of the dead-end complex of E. cloacae MurA with terreic acid.
(Top, stereoview) The overall structure of MurA inactivated by terreic acid exists in an open conformation and is free of UNAG. The loop hosting Cys115 is shown in magenta; the covalently bound terreic acid molecule (cyan) is largely solvent–exposed.
(Middle, stereoview) A detailed view of the Cys115-terreic acid adduct reveals multiple polar interactions (d ≤ 3.3 Å, black dotted lines) with water molecules (orange spheres) and two Ca2+ ions from the crystallization solution (green spheres); hydrophobic interactions (d ≤ 3.8 Å, green dotted lines) exist with Leu138. The blue-colored mesh represents the Fo-Fc difference electron density map (at 2.25 Å resolution and contoured at 3σ), omitting the modified Cys115 residue during the refinement.
(Bottom) Proposed chemical reaction of terreic acid with Cys115.