Figure 1.

Behavioral tasks. A, Center-out task for saccades involved the monkey touching a central target while making a saccade from an adjacent location to one of eight peripheral targets arranged on a square spaced 10°. The lower target is not shown for clarity. B, Center-out task for reaches involved the monkey fixating a central target while making a reach from an adjacent location. C, Saccade relative position-coding task. A saccade is made from one of four initial gaze positions on a line to one of four target positions while a touch is maintained at one of four hand positions on a touch screen. Hand positions and reach targets are shown in green, and gaze positions and saccade targets are shown in red.