Expression of subplate, layer 6, and layer 5 marker genes in Tbr1 null cortex. (A–I) SP and layer 6 markers Ctgf (A–C), Sox5 (D–F), and Tle4 (G–I) had decreased expression in Tbr1 null cortex by ISH and microarray. (J–R) Layer 5 markers Fezf2 (J–L), Er81 (M–O), and Crim1 (P–R) had increased expression. [Scale bars (A–Q): 200 μm.]