TABLE 3.

Coefficient of directionality, the proportion of directional epistasis in total epistasis and the epistatic factor

Linear scale					Cubic scale
Trait	ɛ	(%)		(%)	Trait	ɛ	(%)		(%)
Femur lt	−7.46	40.5	0.937	2.4	(Femur lt)3	−1.92	32.2	0.958	3.1
Humerus lt	−6.59	26.9	0.954	0.7	(Humerus lt)3	−1.46	18.3	0.968	0.6
Tibia lt	−8.45	41.5	0.902	0.0	(Tibia lt)3	−1.73	26.4	0.927	0.0
Ulna lt	−4.58	18.3	0.924	0.7	(Ulna lt)3	−0.78	9.7	0.940	0.6
Tail lt	−2.02	22.6	0.881	1.0	(Tail lt)3	0.19	6.1	0.920	2.0
	−9.90	40.9	0.892	10.1	Body wt	−2.49	31.9	0.904	10.8
	−14.88	16.3	0.896	15.5	Heart wt	−5.47	17.1	0.823	16.2
	−6.01	18.8	0.923	6.9	Kidney wt	−1.12	11.2	0.929	7.6
	−8.33	30.9	0.960	13.5	Spleen wt	−1.88	18.3	0.874	15.5
	−7.72	33.7	0.870	6.4	Liver wt	−1.96	25.7	0.931	7.1

The values of the composite coefficient of directionality (ɛ), ratio between directional and total epistasis , the geometric mean of the absolute epistasis factor (), and the percentage of interactions with negative f (sign epistasis). Results are shown for all traits, on linear and cubic scale. wt, weight; lt, length.