Table 2. Environmental variables.

Name	Description	Hypotheses
HABITAT
Slope	Slope angle of the terrain (degrees)	Griffon vulture is a cliff-nesting raptor, so areas with higher slope will have more cliffs and therefore higher habitat suitability
Elevation	Altitude of the terrain (m.a.s.l.)	Lower elevation can provide protection against inclement weather
Aspect	Terrain exposure classified into eigth categories: north (0°–22.5°; 337.5°–360°),northwest (292.5°–337.5°), west (247.5°–292.5°), southwest (202.5°–245.5°), south (157.5°–202.5°), southeast (112.5°–157.5°), east (67.5°–112.5°) and northeast (22.5°–67.5°)	S or E exposures can provide protection against inclement weather
Topography3	Difference between elevation of the cell and the mean of those included in a moving window of 3×3 side	Ruff terrain can mean more rocky cliffs for nesting but can also increase energy costs of movement
Topography5	Difference between elevation of the cell and the mean of those included in a moving window of 5×5 side
Coverage	Surface coverage according to vulture accesibility: villages (0), forests (1), shrub (2), pasture (3) and rock (4)	Open areas facilitate carcass detection and access
Inhabitants *	Density of inhabitants	Human presence can increase disturbance to breeding colonies but can also provide predictable sources of food
FOOD
LU *	Density of livestock units 1 cow  = 5 livestock units; 1 sheep or goat = 1 livestock unit	Livestock is an important food resource for the species
CowLU *	Density of livestock units of cow
SheepLU *	Density of livestock units of sheep and goat
RedDeer *	Density of captures of red deer	Game species are an important food resource for the species
WildBoar *	Density of captures of wild boar

Environmental variables used to model griffon vulture distribution in the study area.

*Variables calculated at three scales: 90 m-pixel, 3.5 km radius and 10 km radius, centred at the breeding colony.