The optimum, or evolutionary stable, gradient detection strategy ωgi for a solitary individual. (A) Comparison between numerical simulations and analytical calculations of migratory benefits, b, gained by a solitary individual (SI Appendix B). (B) Cost of gradient detection is given by cg = pg(exp(ωg/ωgc) − 1) and (C) comparison between numerical simulations and analytical calculations of the individual fitness (f = b − cg). Parameters: size of population N = 1, pg = 0.75, ωgc = 4.0, strength of noise in perception σr = 0.1.