Figure 4. Different morphologies when optimizing for wiring-constraints only.

We optimized morphologies without the terms for wide-field motion integration, once with only the overall size as constraint and once with the overall size and average path length as constraint. The morphology with the lowest overall size (A) and combined size and path length (B) are shown. The morphology in A deviates substantially from the morphologies found originally, while the morphology in B shares the same blueprint as originally optimized models and VS cells. Both neurons have a significantly lower performance as wide-field motion integrators than the originally optimized passive model neurons.