Table 1.
Protein | Class | Localization | Ca2+-binding | Special properties | Function | References |
---|---|---|---|---|---|---|
Syt 1 | A | Synaptic vesicles LDCVs | Yes | N-terminal N-glycosylation Phospholipids and SNARE binding | Ca2+-sensor for fast exocytosis | 6,10,12,31-33, 39-43,74 |
Syt2 | A | Synaptic vesicles LDCVs? | Yes | N-terminal N-glycosylation Phospholipid and SNARE binding | Ca2+-sensor for fast exocytosis | 8,9,31-33,43, 44,54,74 |
Syt3 | C | Plasma membrane | Yes | Disulfide bonds at N-terminus Phospholipid binding | Ca2+-sensor for exocytosis? | 31-33,74 |
Syt4 | D | Postsynaptic? | No* | Aspartate to Serine substitution in C2A domain loops | unknown*; not essential for survival | 73,74 |
Syt5 | C | Plasma membrane | Yes | Disulfide bonds at N-terminus Phospholipid bindings | Ca2+-sensor for exocytosis? | 31-33,74 |
Syt6 | C | Plasma membrane | Yes | Disulfide bonds at N-terminus Phospholipid binding | Ca2+-sensor for exocytosis? | 31,74 |
Syt7 | B | LDCVs Synapses? | Yes | Multiple splicing isoforms Phospholipid and SNARE binding | Ca2+-sensor for LDCV exocytosis | 47-50,74 |
Syt8 | F | primarily GIT tract | No | Only Syt isoform that is expressed at highest levels outside of brain | Not known | 31,74 |
Syt9 | A | Synaptic vesicles | Yes | Phospholipid and SNARE binding | Ca2+-sensor for fast exocytosis | 43,74 |
Syt10 | C | Plasma membrane | Yes | Disulfide bonds at N-terminus Phospholipid binding | Ca2+-sensor for exocytosis? | 31,74 |
Syt11 | D | Unknown | No | Similar to Syt4, but more abundant | unknown | 74 |
Syt12 | F | Synaptic vesicles | No | Phosphorelated by PKA | May regulate miniature release | 31,74,75 |
Syt13 | F | Not known | No | Not known | 31,74 | |
Syt14 | E | Not known | No | Syt14-16 form a group of related and evolutionarily conserved isoforms | Not known | 76 |
Syt15 | E | Not known | No | Syt14-16 form a group of related and evolutionarily conserved isoforms | Not known | 76 |
Syt16 | E | Not known | No | Syt14-16 form a group of related and evolutionarily conserved isoforms | Not known | 76 |
Synaptotagmins are defined by the presence of an N-terminal transmembrane region, and two C-terminal C2-domains. Each synaptotagmin protein is encoded by a different gene; some are differentially spliced (e.g., Syt7). Classes of synaptotagmins are proposed defined by sequence homologies, conservation, and functions
Syt4 binds to Ca2+ in Drosophila and has been implicated in postsynaptic exocytosis.