TABLE 2.

Parameterization

Symbol	Description	Definition
ρL	Frequency of variable L	e + f + g + h
ρC	Frequency of allele C	c + d + g + h
ρN	Frequency of allele N	b + d + f + h
DNC	Disequilibrium between N and C	d + h − ρNρC
DNL	Disequilibrium between N and L	f + h − ρNρL
DLC	Disequilibrium between L and C	g + h − ρNρC
DNLC	Three-locus disequilibrium	h − ρNρLρC

Reparameterizing the model in terms of the frequencies of individual factors and the disequilibrium between them facilitates biological understanding of what creates associations between factors and phenotypes.