A multiple-reserve DEB model including maintenance, growth, maturation, maturity maintenance and reproduction. Synthesizing units (SUs) control assimilation and allocation to growth, maintenance and reproduction. pA, lumped assimilated material; κP, protein fraction of assimilate; pC, mobilization rate; κM, fraction to growth/maintenance; pS, somatic maintenance; pG, growth allocation; pM, maturity maintenance; pR, reproduction allocation; κR, fraction returned to reserves.