Nutritional niche space calculated for the copepod Acartia tonsa with a DEB model incorporating separate reserves for protein and non-protein assimilates and represented using the geometric framework. (a,b) Egg production, (c,d) lifetime egg production (egg production rate × longevity) without and with (e,f) costs imposed for storage of excess nutrients are presented for eggs with two different C : N ratios. The Redfield ratio is represented as a nutritional rail, with interpolated regions of the state space emphasized relative to the extrapolated regions. See text for further details.