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. 2010 Apr 28;277(1694):2727–2735. doi: 10.1098/rspb.2010.0409

Table 1.

Relatedness (R) between grandparents and grandchildren for the X, Y and autosomes (A). The first term in each cell (fraction or zero) is the level of relatedness assuming no extra-pair paternity and the second term is the level of paternity assurance (Pa = the average probability that no extra-pair paternity occurs in each generation of a line of descent, 0 ≤ Pa ≤1). Figures in bold are explained in the text.

MGM MGF PGM PGF
grandson A 1/4 * 1 1/4 * Pa 1/4 * Pa 1/4 * Pa2
X 1/4 * 1 1/2 * Pa 0 * Pa 0 * Pa2
Y 0 * 1 0 * Pa 0 * Pa 1 * Pa2
granddaughter A 1/4 * 1 1/4 * Pa 1/4 * Pa 1/4 * Pa2
X 1/4 * 1 1/2 * Pa 1/2 * Pa 0 * Pa2
Y 0 * 1 0 * Pa 0 * Pa 0 * Pa2
average over the A 1/4 1/4 * Pa 1/4 * Pa 1/4 * Pa2
 sex of grandchild X 1/4 1/2 * Pa 1/4 * Pa 0
Y 0 0 0 1/2 * Pa2
average over the A 1/8 * (1 + Pa)
1/8 * (1 + Pa)*Pa
 sex of grandchild X 1/8 * (1 + 2Pa)
1/8 * Pa
 and grandparents Y 0
1/4 * Pa2